If you can trust it, Wikipedia sez
this about lower case letters.
Could provide a bit of info for some further investigation. This I see as the most important bit: "Originally alphabets were written entirely in majuscule (capital) letters which were spaced between well-defined upper and lower bounds. When written quickly with a pen, these tended to rounder and simpler forms, like uncials. It is from these that the first minuscule hands developed, the half-uncials and cursive minuscule, which no longer stay bound between a pair of lines.
These in turn formed the foundations for carolingian minuscule, developed by Alcuin for use in the court of Charlemagne, which quickly spread across Europe. Here for the first time it became common to mix both majuscule and minuscule letters in a single text."
(from the Minuscule article on Wikipedia, the free encyclopedia)
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